necessary to study over what scale the phenomenon holds. For example, departure decisions from inflorescences may be affected. Vertical inflorescences sometimes have more nectar in the bottom male flowers than in the upper female flowers (Waddington 1981). Bees tend to arrive at the inflorescence on a lower flower and then move upward (Pyke 1979). Perhaps a bee's weighted assessment of decreasing nectar volume as it moves upward on single inflorescences would hasten its departure for another inflorescence.
Little is known about bees' assessments of pollen quality and quantity and choice (Rasheed & Harder 1997a, b). The amount of pollen available in individual flowers can be detected by bumble bees (Buchmann & Cane 1989; Harder 1990). Waddington etal. (1998) checked to see if the dance rate varied with pollen quality as it does with nectar quality. Pollen loads were collected from honeybees, dried, and ground to a powder. The pollen was given to bees in a petri dish in two ways: pure pollen or pollen mixed with alpha-cellulose powder (1:1 by volume). Alpha-cellulose powder does not have nutritional value to bees; thus, the mixture is lower in quality than pure pollen. The dance rate was higher after bees
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