Species contend effectively with diminishing, or increasingly distant nutrient

resources, either through exploiting alternative sources (nitrogen fixation, phos-

phatase production, phagotrophy on bacteria or particulate organic material).

Most are motile, some are strongly so, sinking rates low but may undertake con

trolled migrations over large vertical distances. Also referred to as resource 'glean

ers' (Anderies and Beisner 2000). Representative genera Microcystis, Anabaena,

Gloeotrichia, Ceratium, Peridinium, Chrysosphaerella, Uroglena.

SS strategists

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