Many factors influence the spatial distribution of a species, and it is difficult to determine whether the arrangement of individuals in a habitat is determined by one, a few, or the combined action of all of them. Individuals may move in response to temporal changes (daily rhythms, weather, season), or to fulfill varying needs (dispersal, mate finding, etc.) (Basset et al., 2003b). The distribution of cockroach individuals is often correlated with the proximity of appropriate food sources. In sparsely vegetated sites, for example, cockroaches are frequently associated with whatever plants (and therefore their litter) are present. This includes deserts (Edney et al., 1978), alpine zones (Sinclair et al., 2001), and other arid or Mediterranean-type habitats such as southwestern Australia, where the number and diversity of ground-dwelling cockroaches depends on the type, percent cover, and depth of the litter present (Abenserg-Traun et al.,
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