The record of trace fossils is unfortunately extremely impoverished. Well-attested arthropod trace fossils from the Cambrian, such as Cruziana and Rusophycus, are normally assigned to the trilobites (Hughes, this volume; but see also Pratt 1994; Crimes, this volume). Other traces may well also have an arthropodan origin, but evidence based on trace and trace maker co-occurrence and functional morphology is lacking. A notable exception is provided by traces from the Czech Paseky Shale, which are attributed to various nontrilobite arthropods and appear to have been made in a nonmarine environment (Chlupac 1995; Mikulas 1995; see also Osgood 1970 and Hes-selbo 1988 for examples of aglaspidid traces). Traces that can be confidently assigned to chasmataspid chelicerates are known from the Upper Cambrian of Texas (Dunlop et al. 1996). Finally, there is a limited amount of information from the study of copro-lites (e.g., those attributed to Anomalocaris by Conway Morris and Robison ).
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