Most of the work described here was funded by NERC Grant NER/A/S/2002/00882 and BBSRC Grant 43/P19284. We thank Lynne Boddy for detailed protocols for soil-based microcosms, and the P. velutina isolate used in these experiments. Strains S7 and S16 of S. lacrymans were provided by Olaf Schmidt, University of Hamburg. Dan Eastwood (University of Warwick) works with Monika Tlalka on gene expression in P. velutina. We are grateful for photography by John Baker, and technical assistance from Caroline O'Brien. Our thanks to Mark Fricker for helpful discussion.

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