Funding from The Swedish Research Council for Environment, Agricultural Sciences and Spatial Planning (FORMAS) as well as The Swedish University of Agricultural Sciences (SLU) is gratefully acknowledged. Thanks are also due to Dr. Katarina Ihrmark and Professors Roger Finlay, Jan Stenlid, Peter Hogberg, Torgny Nasholm, Tryggve Persson and Bjorn Berg, for research collaboration and Dr. Andy Taylor, Dr. Petra Fransson and Professor Lynne Boddy and Dr. Juliet Frankland, for critical reading of the manuscript.

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