Mallada basalis (Walker), distributed widely across the Indo-Pacific region, produces audible buzzing noises by striking the modified costal margins of its vibrating hind wings forcibly against the substrate (Duelli and Johnson, 1982). Only the male has the thickened, hammer-like pterostigma on the wing required for signalling. Males call vigorously in the presence of females, and are capable of producing the loudest airborne sounds of any neuropterid.
Percussive sounds can also be found in lacewings which tremulate, e.g. two species in the carnea group of Chrysoperla (Henry et al., 2002, 2003). In such cases, the tremulating abdomen will periodically strike the substrate, usually toward the end of a song consisting of a single long volley (Chrysoperla agilis; Henry et al.) or many short volleys (C. pallida; Henry et al.). The sound which we hear is a low-intensity ticking or rattling noise, detectable over a range of perhaps 25 cm.
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