The lacunarity is a measure of spatial arrangement between two or more patches. This index focuses on the empty spaces between patches and is particularly sensitive to their spatial arrangement. In particular lacunarity is important for processes or species patterns that happen outside the habitat patches, like dispersion, predation, and settlement. Lacunarity measures the pioneering capacity of species, or the environmental pressure of the more or less hostile environment on a species that is in movement from one habitat patch to another.
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