Bioenergetic models describe the factors affecting the growth of an individual over its lifetime. The rate of change in average individual biomass (B ) is modeled as the sum of weight-specific rates of physiological processes: consumption (C), respiration (R '), egestion (F), excretion (U), and reproductive loss (G '):
dB ',,,,,,, — — C ' - (R ' + F ' + U ' + G' ) 
These process rates in turn are considered functions of other variables, such as water temperature or food availability. A population model based on bioenergetics must also consider mortality and recruitment, two forces not included in eqn .
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