Litter collection

Using traps, in nets, or sampled on the ground is useful for LAI calculation of broadleaved plants. Species composition in the canopy is not distinguished when using canopy LAI. By sorting leaves from different species in mixed broadleaved forest, the contribution of each species from the community may be quantified (Figure 5). Few data exist on leaf biomass of temperate deciduous forest communities which include undergrowth. If biomass is nearly negligible as compared to total stand biomass, the

■ Fagus □ Quercus □ Betula □ Carpinus Fraxinus □ Prunus

■ Fagus □ Quercus □ Betula □ Carpinus Fraxinus □ Prunus

20 32 36 96 99 102 114 116 167

Figure 5 Species contribution to total stand LAI in beech-dominated forest as estimated by litter collection and species sorting. Each plot number is related to spatial variability within the stand. For further details, see Bouriaud O, Soudani K, and Breda N (2003) Leaf area index from litter collection: Impact of specific leaf area variability within a beech stand. Canadian Journal of Remote Sensing 29: 371-380.

20 32 36 96 99 102 114 116 167

Figure 5 Species contribution to total stand LAI in beech-dominated forest as estimated by litter collection and species sorting. Each plot number is related to spatial variability within the stand. For further details, see Bouriaud O, Soudani K, and Breda N (2003) Leaf area index from litter collection: Impact of specific leaf area variability within a beech stand. Canadian Journal of Remote Sensing 29: 371-380.

contribution of leaf area of undergrowth has been demonstrated to be a significant part (from 15% to 60%) of LAI of the whole community (tree or shrub layer + ground vegetation) within a narrow range of 7. Finally, the measured leaf properties (individual leaf area, specific leaf area, number of leaves, etc.) are of key importance for a comprehensive analysis of LAI changes between stands, species, or dates. Indeed, differences in LAI for a given stand during certain years could be the result of any change in number of plants, number of leaves, or individual leaf area.

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