The interpretation of these indices is discussed in depth in the next paragraphs; for now, it is sufficient to say that the total capacity C represents the maximum potential that a system has at its disposal to achieve further development and it serves as upper boundary for ecosystem organization. The capacity is then partitioned into flows organization (ascendency A) and redundant, unorganized flows (overhead $).
These measures have been extended to introduce the biomass (standing stock) of compartments into the computation. In fact, we expect the fluxes between compartments to be highly dependent on the availability of biomass (standing stock). Moreover, standing stock represents the most reliable source of empirical data, while the fluxes themselves are often obtained from estimates that are highly dependent on the biomasses. For example, the formula for ascendency, given that Bj is the biomass of compartment jand B is the total biomass in the system, becomes (using Kullback-Leibler formulation)
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