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Humus-enzyme complexes ^ Clay-enzyme complexes ^

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Activities persist in harsh conditions inhibiting microbial activity

Intracellular enzyme activities Fauna Roots

Intracellular enzyme activities Fauna Roots

Respond to changes in microbiological activity

Microbial functional diversity

Fungi Bacteria

FIGURE 3.8. Various activities contributing to the overall enzyme activity measured in soil compared with those affecting the microbial functional diversity in soil (Nannipieri et al., 2002).

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