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The right of Mike Begon, Colin Townsend and John Harper to be identified as the Authors of this Work has been asserted in accordance with the UK Copyright, Designs and Patents Act 1988.
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First edition published 1986 by Blackwell Publishing Ltd Second edition published 1990 Third edition published 1996 Fourth edition published 2006
Library of Congress Cataloging-in-Publication Data Begon, Michael.
Ecology : from individuals to ecosystems / Michael Begon, Colin R. Townsend, John L. Harper.—4th ed. p. cm.
Includes bibliographical references and index. ISBN-13: 978-1-4051-1117-1 (hard cover : alk. paper) ISBN-10: 1-4051-1117-8 (hard cover : alk. paper)
1. Ecology. I. Townsend, Colin R. II. Harper, John L. III. Title. QH54.B416 2005
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