For the redundancy and the cost of information we get correspondingly: R2 = 0.265 and C2 = 1.36. Since at this level the elementary unit is a two-letter word, the information per letter will be equal to /2 < 3.6 bits.

Note that in addition to their area, the biomes are also characterised by three values: annual productivity Pi, living biomass Bi, and dead organic matter Di (see Table 4.2). It is obvious that we can define the frequencies pj, pf and pf for each value as pj = PiS/X301 PiS; pf = BiS/X3°1 BiS and pf = Ds/X301 DSi. The description in terms of productivity, living or dead organic matter can be considered as description at the first level with information per letter defined as

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